This paper was a model of clarity, Richard told his students. It represented everything that science should be. Richard paced in front of the chalkboard, speaking easily and without notes. Like the minor evolutionist Robert Chambers, he had been born hexadactylic; he was sensitive about his left hand, which was somewhat scarred from the childhood operation that had removed his extra finger. And so, although Richard gestured freely he used only his right hand and kept his left in his pocket. From the back of the room, where I sat when I came each fall to hear him lecture, I could watch the students listen to him. After he passed out the paper, Richard told the students his first, conventional version of Gregor Mendel’s life. Mendel, he said, grew up in a tiny village in the northwestern corner of Moravia, which was then a part of the Hapsburg Empire and later became part of Czechoslovakia. When he was twenty-one, poor and desperate for further education, he entered the Augustinian monastery in the capital city of Brünn, which is now called Brno.